古鲸

古鲸小目 化石时期：
巴基鲸（上）与走鲸（下）的骨骼模拟
科学分类
界：	动物界 Animalia
门：	脊索动物门 Chordata
纲：	哺乳纲 Mammalia
目：	偶蹄目 Artiodactyla
下目：	鲸下目 Cetacea
并系群：	†古鲸类 Archaeoceti Flower, 1833
传统分类中包含的子类群
†巴基鲸科 Pakicetidae †步鲸科 Ambulocetidae †雷明顿鲸科 Remingtonocetidae †原鲸科 Protocetidae †龙王鲸科 Basilosauridae †矛齿鲸科 Dorudontidae
传统分类中未包含，但在 演化层面上包含的演化支
须鲸小目 Mysticeti 齿鲸小目 Odontoceti

辛西娅鲸（左）与走鲸（右）的骨骼

东京国立科学博物馆展示的库奇鲸骨骼

华圣顿史密森尼博物馆展示的轭根鲸骨骼

撒格哈鲸的骨骼

古鲸小目（学名：Archaeoceti）也作原鲸小目，即古代鲸鱼，包括现代鲸鱼的祖先。古鲸小目是鲸下目三个小目之一，传统上用以概括史前鲸豚类，但不包括其存活至现代的后代，因此，这是一个并系群分类，不是自然的演化支。古鲸拥有后肢，不像今日的鲸类。最早的古鲸可能完全在陆地上活动。其灭绝原因至今众说纷纭，其中一种解释是由于气候变化，海洋变得过份寒冷，造成古鲸类灭绝。

分类

• 古鲸小目 Archaeoceti Flower, 1833
  • 巴基鲸科 Pakicetidae Gingerich e Russell, 1990
    • 巴基鲸 Pakicetus Gingerich e Russell, 1981
    • 娜拉鲸 Nalacetus Thewissen e Hussain, 1998
    • 鱼中兽 Ichthyolestes Dehm e Oettingen-Spilberg, 1958
  • 步鲸科 Ambulocetidae Thewissen, Madar e Hussain, 1996
    • 走鲸 Ambulocetus Thewissen, Hussain e Arif, 1994
    • 甘达克鲸 Gandakasia Dehm e Oettingen-Spilberg, 1958
    • 喜马拉雅鲸 Himalayacetus Bajpai e Gingerich, 1998
  • 雷明顿鲸科 Remingtonocetidae Kumar e Sahni, 1986
    • 安德鲁斯鲸 Andrewsiphius Sahni e Mishra, 1975
    • 雷明顿鲸 Remingtonocetus Kumar e Sahni, 1986
    • 大连特鲸 Dalanistes Gingerich, Arif e Clyde, 1995
    • 阿托克鲸 Attockicetus Thewissen e Hussain, 2000
    • 库奇鲸 Kutchicetus Bajpai e Thewissen, 2000
    • Rayanistes Bebej, Zalmout, El-Aziz, Antar, and Gingerich, 2016
  • 原鲸科 Protocetidae Stromer, 1908
    • 埃及鲸属 Aegyptocetus Bianucci & Gingerich 2011
      • 埃及鲸 Aegyptocetus tarfa Bianucci & Gingerich 2011
    • Tupelocetus
    • 原鲸亚科 Protocetinae Gingerich et al. 2005)
      • 熊神鲸 Artiocetus Gingerich, ul Haq, Zalmout, Khan & Malkani, 2001
      • 圆齿鲸 Crenatocetus McLeod & Barnes 2008
      • Dhedacetus Bajpai and Thewissen, 2014
      • 加氏鲸 Gaviacetus Gingerich, Arif e Clye, 1995
      • 印支鲸 Indocetus Sahni e Mishra, 1975
      • Kharodacetus Bajpai and Thewissen, 2014
      • 慈母鲸 Maiacetus Gingerich et al., 2009
      • 原鲸 Protocetus Fraas, 1904
      • 乔丹斯鲸 Qaisracetus Gingerich,Ul-Haq, Khan e Zalmout, 2001
      • 罗德侯鲸 Rodhocetus Gingerich, Raza, Arif, Anwar e Zhou, 1994
      • 泰卡鲸 Takracetus Gingerich, Arif e Clyde, 1995
      • 多哥鲸 Togocetus Gingerich & Cappetta 2014
      • Peregocetus Lambert et al., 2019
    • 麦卡鲸亚科 Makaracetinae P. D. Gingerich et al. 2005
      • 麦卡鲸 Makaracetus P. D. Gingerich et al. 2005
    • 乔治亚鲸亚科 Georgiacetinae Gingerich et al., 2005
      • 巴比亚鲸 Babiacetus Trivedy e Satsangi, 1984
      • 卡罗莱鲸 CarolinacetusGeisler et al. 2005
      • 始原鲸 Eocetus Fraas, 1904
      • 乔治亚鲸 Georgiacetus Hulbert, Petkewich, Bishop, Bukry e Aleshire, 1998
      • 那支特鲸 Natchitochia Uhen, 1998
      • 帕坡氏鲸 Pappocetus Andrews, 1920
      • Pontobasileus Leidy 1873
  • 龙王鲸科 Basilosauridae Cope, 1868
    • 龙王鲸亚科 Basilosaurinae Cope, 1868
      • 帝王鲸 Basilotritus Goldin & Zvonok 2013
      • 龙王鲸 Basilosaurus Harlan, 1834
      • 王鲸 Basiloterus Gingerich, Arif, Bhatti, Anwar e Sanders, 1997
      • Eocetus Fraas, 1904
      • Platyosphys Kellogg, 1936
  • 矛齿鲸科 Dorudontidae (Miller, 1923)
    • 矛齿鲸亚科 Dorudontinae Miller, 1923
      • 弯臂鲸 Ancalecetus Gingerich e Uhen, 1996
      • 金原鲸 Chrysocetus Uhen e Gingerich, 2001
      • 辛西娅鲸 Cynthiacetus Uhen, 2005
      • 矛齿鲸 Dorudon Gibbes, 1845
      • 奥库卡赫鲸 Ocucajea Uhen et al. 2011
      • 海乡鲸 Pontogeneus Leidy, 1852
      • 施特罗默鲸 Stromerius Gingerich, 2007
      • 撒格哈鲸属 Saghacetus Gingerich, 1992
      • 死神鲸 Supayacetus Uhen et al. 2011
      • 轭根鲸 Zygorhiza True, 1904

种系发生学

鲸形类   †劳氏兽科 Raoellidae   鲸下目   †巴基鲸科 Pakicetidae       †步鲸科 Ambulocetidae       †雷明顿鲸科 Remingtonocetidae       †原鲸科 Protocetidae       †龙王鲸科 Basilosauridae       †矛齿鲸科 Dorudontidae       齿鲸小目 Odontoceti     须鲸小目 Mysticeti               Cetacea Cetaceomorpha

†古鲸小目 Archaeoceti

地质时期

各类古鲸的化石所在地质时期

参考文献

• Andrews, C. W. A description of new species of zeuglodont and of leathery turtle from the Eocene of Southern Nigeria (PDF). Proceedings of the Zoological Society of London 1919. 1920, 18: 309–19 [April 2013]. OCLC 228169273. （原始内容 (PDF)存档于2015-01-04）. 
• Bajpai, Sunil; Gingerich, Philip D. A new Eocene archaeocete (Mammalia, Cetacea) from India and the time of origin of whales. PNAS. December 1998, 95 (26): 15464–68. OCLC 678707438. doi:10.1073/pnas.95.26.15464. 
• Bajpai, S; Thewissen, JG; Sahni, A. The origin and early evolution of whales: macroevolution documented on the Indian subcontinent (PDF). J Biosci. November 2009, 34 (5): 673–86 [February 2013]. OCLC 565869881. PMID 20009264. doi:10.1007/s12038-009-0060-0. 
• Dehm, Richard; Oettingen-Spielberg, Therese zu. Paläontologische und geologische Untersuchungen im Tertiär von Pakistan. 2. Die mitteleocänen Säugetiere von Ganda Kas bei Basal in Nordwest-Pakistan. Abhandlungen / Neue Folge, 91. Munich: Beck. 1958. OCLC 163296508. 
• Flower, William Henry. On the Arrangement of the Orders and Families of existing Mammalia (PDF). Proceedings of the Zoological Society of London. 1883: 178–86. OCLC 83091701. 
• Fordyce, Ewan. Cetacean Evolution. Perrin, William R; Wiirsig, Bernd; Thewissen, J G M (编). Encyclopedia of Marine Mammals. Academic Press. 2002: 214–25. ISBN 0-12-551340-2. 
• Fraas, E. Neue Zeuglodonten aus dem unteren Mitteleozän von Mokattam bei Cairo 6 (3). Jena: Verlag Gustav Fischer: 199–220. 1904. 
• Geisler, Jonathan H; Sanders, Albert E; Luo, Zhe-Xi. A new protocetid whale (Cetacea, Archaeoceti) from the late middle Eocene of South Carolina. American Museum Novitates. July 2005, 3480: 1–68. OCLC 4630500767. doi:10.1206/0003-0082(2005)480[0001:ANPWCA]2.0.CO;2. 
• Gingerich, Philip D; Haq, U; Khan, H; Zalmout, S. Eocene stratigraphy and archaeocete whales (Mammalia, Cetacea) of Drug Lahar in the eastern Sulaiman range, Balochistan (Pakistan). Contributions from the Museum of Paleontology, University of Michigan. 2001, 30 (11): 269–319. OCLC 50061585. 
• Gingerich, Philip D; Haq, U; Zalmout, S; Khan, H; Malkani, S. Origin of whales from early artiodactyls: hands and feet of Eocene Protocetidae from Pakistan. Science. September 2001, 293 (5538): 2239–2242. Bibcode:2001Sci...293.2239G. ISSN 0036-8075. PMID 11567134. doi:10.1126/science.1063902. 
• Gingerich, Philip D; Arif, M; Bhatti, M Akram; Anwar, M; Sanders, William J. Basilosaurus drazindai and Basiloterus hussaini, New Archaeoceti (Mammalia, Cetacea) from the Middle Eocene Drazinda Formation, with a Revised Interpretation of Ages of Whale-Bearing Strata in the Kirthar Group of the Sulaiman Range, Punjab (Pakistan). Contributions from the Museum of Paleontology, University of Michigan. 1997, 30 (2): 55–81. OCLC 742731913. 
• Gingerich, Philip D. Stromerius nidensis, new archaeocete (Mammalia, Cetacea) from the Upper Eocene Qasr El-Sagha Formation, Fayum, Egypt (PDF). Contributions from the Museum of Paleontology. 2007, 31 (13): 363–78. OCLC 214233870. 
• Gingerich, Philip D.; Arif, Muhammad; Clyde, William C. New Archaeocetes (Mammalia, Cetacea) from the middle Eocene Domanda Formation of the Sulaiman Range, Punjab (Pakistan). Contributions from Museum of Paleontology, The University of Michigan. 1995, 29 (11): 291–330 [February 2013]. OCLC 34123868. 
• Gingerich, Philip D.; Raza, S. M.; Arif, M.; Anwar, M.; Zhou, X. New whale from the Eocene of Pakistan and the origin of cetacean swimming. Nature. 1994, 368: 844–47 [April 2013]. OCLC 742745707. doi:10.1038/368844a0. 
• Gingerich, Philip D.; Russell, Donald E. Pakicetus inachus, A New Archaeocete (Mammalia, Cetacea) from the Early-Middle Eocene Kuldana Formation of Kohat (Pakistan) (PDF). Contributions from the Museum of Paleontology, The Museum of Michigan. 1981, 25 (11) [February 2013]. OCLC 742729300. 
• Gingerich, Philip D.; ul-Haq, Munir; von Koenigswald, Wighart; Sanders, William J.; Smith, B. Holly; Zalmout, Iyad S. New Protocetid Whale from the Middle Eocene of Pakistan: Birth on Land, Precocial Development, and Sexual Dimorphism. PLoS ONE. 2009, 4 (2): e4366. OCLC 678622523. PMC 2629576 . doi:10.1371/journal.pone.0004366. 
• Gingerich, Philip D.; Zalmout, Iyad S.; Ul-Haq, Munir; Bhatti, M. Akram. Makaracetus bidens, a new protocetid archaeocete (Mammalia, Cetacea) from the early middle Eocene of Balochistan (Pakistan) (PDF). Contributions from the Museum of Paleontology. 2005, 31 (9): 197–210 [March 2013]. OCLC 742723177. 
• Hulbert, Jr, Richard C; Petkewich, Richard M; Bishop, Gale A; Bukry, David; Aleshire, David P. A New Middle Eocene Protocetid Whale (Mammalia: Cetacea: Archaeoceti) and Associated Biota from Georgia. Journal of Paleontology. September 1998, 72 (5): 907–927. JSTOR 1306667. OCLC 4908698029. 
• Kumar, K.; Sahni, A. Remingtonocetus harudiensis, new combination, a middle Eocene archaeocete (Mammalia, Cetacea) from western Kutch, India. Journal of Vertebrate Paleontology. 1986, 6 (4): 326–349. OCLC 4649653943. doi:10.1080/02724634.1986.10011629. 
• McLeod, S. A.; Barnes, L. G. A new genus and species of Eocene protocetid archaeocete whale (Mammalia, Cetacea) from the Atlantic Coastal plain. Wang, Xiaoming; Barnes, Lawrence G. (编). Geology and Vertebrate Paleontology of Western and Southern North America (PDF) 41. 2008: 73–98 [February 2013]. 
• Rose, Kenneth David. The Beginning of the Age of Mammals. JHU Press. 2006. ISBN 978-0-8018-8472-6. 
• Sahni, Ashok; Mishra, Vijay Prakash. Lower Tertiary vertebrates from western India. Monograph of the PaleontologicalSociety of India. 1975, 3: 1–48. ASIN B0007AL8UE. OCLC 3566369. 
• Thewissen, JGM. Archaeocetes, Archaic. Perrin, William R; Wiirsig, Bernd; Thewissen, J G M (编). Encyclopedia of Marine Mammals. Academic Press. 2002: 36–9. ISBN 0-12-551340-2. 
• Thewissen, J.G.M.; Hussain, S.T. Attockicetus praecursor, a new remingtonocetid cetacean from marine Eocene sediments of Pakistan. Journal of Mammalian Evolution. 2000, 7 (3): 133–46. OCLC 362777268. doi:10.1023/A:1009458618729. 
• Thewissen, J. G. M.; Hussain, S. T. Systematic review of the Pakicetidae, Early and middle Eocene Cetacea (Mammalia) from Pakistan and India. Bulletin of the Carnegie Museum. 1998, 34: 220–38. 
• Thewissen, J.G.M.; Madar, S.I.; Hussain, S.T. Ambulocetus natans, an Eocene cetacean (Mammalia) from Pakistan. Courier Forschungsinstitut Senckenberg 191. 1996: 1–86. ISBN 9783929907322. OCLC 36463214. 
• Thewissen, J. G. M.; Williams, E. M.; Roe, L. J.; Hussain, S. T. Skeletons of terrestrial cetaceans and the relationship of whales to artiodactyls (PDF). Nature. 2001, 413: 277–81 [February 2013]. OCLC 118116179. doi:10.1038/35095005. 
• Trivedy, A. N.; Satsangi, P. P. A new archaeocete (whale) from the Eocene of India. Abstracts of 27th International Geological Congress, Moscow. 1984, 1: 322–23. 
• True, F.W. The fossil cetacean, Dorudon serratus GIBBES. Bulletin of the Museum of Comparative Zoology. Bulletin of the Museum of Comparative Zoology. 1908, 52 (4): 5–78. OCLC 355813868. OL 19219818M. 
• Uhen, Mark D. New protocetid (Mammalia, Cetacea) from the late middle Eocene Cook Mountain Formation of Louisiana. Journal of Vertebrate Paleontology. 1998, 18 (3): 664–8. OCLC 204612030. 
• Uhen, Mark D; Gingerich, Philip D. New genus of dorudontine archaeocete (Cetacea) from the middle-to-late Eocene of South Carolina. Marine Mammal Science. January 2001, 17 (1): 1–34. OCLC 204061291. doi:10.1111/j.1748-7692.2001.tb00979.x. 
• West, Robert M. Middle Eocene large mammal assemblage with Tethyan affinities, Ganda Kas region, Pakistan. Journal of Paleontology. 1980, 54 (3): 508–533. JSTOR 1304193. OCLC 4899161959. 

外部链接

• Archaeoceti