拉斯克基礎醫學研究獎、沃爾夫醫學獎、托馬斯·亨特·摩爾根獎章、哈維獎、蓋爾德納國際獎、國際生物學獎、路易莎·格羅斯·霍維茨獎、NAS Award in the Neurosciences、羅森斯特爾獎、Karl Spencer Lashley Award、Bristol-Myers Squibb Award for Distinguished Achievement in Neuroscience Research、Robert J. and Claire Pasarow Foundation Award for Distinguished Contributions to Neuropsychiatric Research、Crafoord Prize in Biosciences、Bower Award and Prize for Achievement in Science、英國皇家學會外籍會員、Ralph W. Gerard Prize、honorary doctor of Purdue University、Mendel Medal、Howard Taylor Ricketts Prize